Puppeteer Craig Schwartz and animal lover and pet store clerk Lotte Schwartz are just going through the motions of their marriage. Despite not being able to earn a living solely through puppeteering, Craig loves his profession as it allows him to inhabit the skin of others. He begins to take the ability to inhabit the skin of others to the next level when he is forced to take a job as a file clerk for the off-kilter LesterCorp, located on the five-foot tall 7½ floor of a Manhattan office building. Behind one of the filing cabinets in his work area, Craig finds a hidden door which he learns is a portal into the mind of

Army Captain Colter Stevens finds himself working on a special program where his consciousness can be inserted into another human being. The only catch is can only be there for 8 minutes at any given time. That morning, a bomb exploded on a commuter train just outside Chicago. He occupies the body of teacher going to work on that train and is confused as to what he is doing or why he is there as his last memory is of flying his helicopter on a combat mission in Afghanistan. Those in charge of the program explain to him that there is a bomb on the train, and that he must locate it. More importantly, he must identify the bomber as another bombing is expected later that day. He is also told however that he cannot change the past and can only gather information. As he develops a liking for his traveling companion Christina, he sets out to test that theory.

Much to his surprise, timid Joel Barish is shocked to discover that the love of his life, sparky Clementine, has had him erased from her memory. To pay her back in the same coin, poor Joel summons up the courage to undergo a painless but intricate medical procedure to do the same, utterly unaware that darkness is an essential part of the light. Now, as hurt and angry Joel's ugly recollections of Clementine gradually fade away, giving way to a soulless black void, suddenly, he begins having second thoughts, toying with the idea of stopping the irreversible process. In the end, is ignorance really bliss?

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